Darwin ____ on the Origin of species probably ranks second o
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选B毫无疑问

整个句子是这样的:Darwin whose On the Origin of Species probably ranks second only to the Bible , had a great impact on western thought.

主句为 Darwin had a great impact on western thought.

定语从句为 whose On the Origin of Species probably ranks second only to the Bible,

因为 On the Origin of Species 是书名《物种起源》,它和Darwin(达尔文)是所有关系,所以用关系词whose

意为:达尔文给西方思想以深远的影响,他的《物种起源》或许是一本仅次于《圣经》的著作.